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Meaning of columella

- An axis to which a carpel of a compound pistil may be attached, as in the case of the geranium; or which is left when a pod opens.
- A columnlike axis in the capsules of mosses.
- A term applied to various columnlike parts; as, the columella, or epipterygoid bone, in the skull of many lizards; the columella of the ear, the bony or cartilaginous rod connecting the tympanic membrane with the internal ear.
- The upright pillar in the axis of most univalve shells.
- The central pillar or axis of the calicles of certain corals.